Program of Study


Graduate Courses

The Graduate Group offers as much flexibility as possible to students as they develop their scientific interests, while emphasizing the development of students into well-rounded neuroscientists. The required first-year core course is designed to provide a survey of the major issues and techniques in modern neuroscience, from molecules to cognition. In their second year, students are encouraged to build on their core neuroscience foundation by taking upper level graduate elective courses and specialty courses offered at other universities. Individual course selections are arranged in consultation with the Graduate Group Master Advisor in the first year, and then with the student's thesis advisor thereafter. Formal course work is usually finished by the end of the second year.

Required Neuroscience Core Curriculum (First Year)

NSC 221 Cellular Neurophysiology
Fall: Lecture--3 hours; Discussion--1 hour. Physiological aspects of cellular and subcellular organization of the nervous system. Neuronal cell biology, the structure and function of ion channels, electrical excitability, signaling cascades, sensory transduction and, mechanisms of synaptic transmission, and the cellular basis of learning and memory. Instructors: Trimmer, Chen
2008-09: T, W, Th, 9:30-11, TBA

NSC 201 Neuroanatomy
Fall: Lecture--2 hours; Lab/Discussion--1 hour. Lectures, demonstrations, and dissections, emphasizing functional significance of neuroanatomy from a biological perspective, with comparisons between human and non-human brains. Emphasis placed on functional anatomy of the nervous system, integrated with cellular, molecular, cognitive, and developmental concepts. Instructors: Jones, Amaral, Chapman, Cheng, Usrey
2008-09: T, W, Th, 1:10-2:00, 320 Medical Neurosciences Building

NSC 222 Systems Neuroscience
Winter: Lecture--4 hours; Discussion--1 hour. Advanced course covering the integrative and information-processing aspects of nervous system organization. Topics include sensory systems, motor function, sensorimotor integration, the limbic system, and the neurobiology of learning and memory. Instructors: Ditterich, DeBello
2008-09: T,Th 9-10:15; W 9-11:30 320 Medical Neurosciences Building


NSC 223 Cognitive Neuroscience

Spring: Lecture--3 hours; Discussion--1 hour. Neurobiological bases of higher mental function including attention, memory, language. Instructors: T. Swaab, G. Mangun
2008-09: TBA, Center for Mind and Brain


NSC 224A Molecular Neuroscience
Winter: Lecture--2 hours. Key issues in developmental and molecular neurobiology. Discussion emphasis on critical evaluation of the experiments and methods described in research papers. Readings of seminal, primary research papers, reviews, and book chapters. Reading materials will be distributed one week in advance. Instructors: Diaz, L'Etoile

2008-09: F, 9:00-11:00, 320 Medical Neurosciences Building

NSC 224B
Developmental Neuroscience
Spring: Lecture--2 hours. Continuation of NSC 224A: Key issues in developmental and molecular neurobiology, focusing on developmental topics. Discussion emphasis on critical evaluation of experiments and methods described in associated literature. Instructors: Chapman, McAllister
2008-09: TBA

Elective Courses Offered though the Neuroscience Graduate Group (Second Year)

NSC 243 - Topics in Cellular and Behavioral Neurobiology
Lecture/ Discussion­1 hour. An advanced examination of several current problems in neurobiology. Topics will vary in different years; may be repeated for credit. Instructor: Ishida

NSC 261A - Topics in Vision
Lecture/discussion -2hours. Visual system functions and underlying mechanisms. Emphasis on recent research and pertinent aspects of anatomy, biochemistry, electrophysiology, psychophysics, development and genetics. Topics may vary each year. Offered every other year. Instructor: Ishida


NSC 298A - Advanced Cellular Neurobiology
Lecture/ Discussion. This course covers selected topics in neurophysiology, from ion channels to simple circuits, for students with a major interest in cellular neuroscience. Topics to be covered include channel biophysics, action potential propagation, intracellular signal transduction pathways, synaptic physiology and quantal analysis, cellular mechanisms of synaptic plasticity, and neuromodulation of synaptic circuitry. Instructors: Burns, Chen, and Mulloney

NSC 220 - Seminar Speaking
Lecture/ Presentations-3 hours. Each student will present a formal, 45-minute seminar (using power point or slides) to members of the Center for Neuroscience. Students are encouraged to present 2 seminars, each in consecutive quarters. Seminar topics are student-selected, and must be discussed with the instructor prior to selection. Offered winter and spring quarters. Instructor: K. McAllister and W. DeBello
2008-09: Winter: TBA; Spring TBA

NSC/NPB 216- Ion Channels
Lecture­3 hours. Graduate level course on properites of ion channels. Offered alternating years. Instructor: Pappone

MED 420- Clinical Neuroscience
Fred Gorin is the new IOR for this course, starting this quarter. This is a good class to learn about different neurological diseases, their etiologies, and their treatments. A variety of lecturers parades through, of varying quality. But most should be good. I highly recommend at least checking out the schedule and seeing if any lectures interest you.

NPB 270. How to Write a Fundable Grant Proposal
Lecture/discussion--3 hours. Prerequisite: graduate standing in a life science and consent of instructor. Familiarization with the skills required to craft a successful grant proposal submitted to extramural agencies such as NIH and NSF. Instructor: Chalupa

NPB 292. Cortical Plasticity and Perception
Lecture/discussion--2 hours. Examination of research articles on cortical plasticity and changes in perception. Examples drawn from studies of the somatosensory, visual, auditory, and motor cortex. Offered in alternate years. Instructor: Recanzone.

NPB 263. Modeling in Systems Neuroscience
Lecture--3 hours; lecture/laboratory--1 hour. Modeling as a tool in systems neuroscience. Mathematical techniques will be introduced and used to explore advanced topics in echolocation, sound localization, electroreception, communications, and motor systems. Other topics include transforms, modeling assumptions, scales and linearity. Offered in alternate years. Instructor: Sutter

NPB 163. Information Processing Models in Neuroscience and Psychology

Lecture--3 hours; term paper. Basic mathematical modeling techniques used in neuroscience and psychology. Specific topics include linear systems theory, Fourier transforms, neural networks, adaptive systems, probabilistic inference and information theory. Emphasis on understanding information processing in neural systems. Instructors: Sutter

Elective Courses Offered by other Departments


Specialty Courses


Students are also encouraged to take specialty courses offered during the summers at a number of sites across the country. These courses are often specialized and expose students to experts in a particular field of research. For example, 4 of our students participated in the "Structure, Function, and Development of the Visual System" 2001 Summer Course at Cold Spring Harbor Labs.

Links to Specialty Courses:

Cold Spring Harbor Laboratory

Marine Biological Laboratory at Woods Hole

Friday Harbor Laboratory

Bodega Marine Laboratory



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